**Supplementary Materials:** Supplementary materials can be found at http://www.mdpi.com/1422-0067/20/22/ 5709/s1.

**Author Contributions:** M.S.S. initiated and designed the experiment. W.Q., R.Y., B.T. and Y.A.H. helped in conducting experiments. M.S.S. performed the RNA extractions and plant hormone analysis. M.S.S., Y.A.H., W.Q., N.U.K. and H.S. (Hiba Shaghaleh) analyzed the data and results. M.S.S. monitored the experimental work and wrote the manuscript. M.S.S. and H.S. (Hongbo Shao) revised and edited the final version of the manuscript. All authors read and approved the final manuscript.

**Funding:** This work was supported by Jiangsu Autonomous Innovation of Agricultural Science & Technology [CX (15)1005], the Excellent Scientist Plan of JAAS, and Shuangchuang Talent Plan of Jiangsu Province.

**Acknowledgments:** We thank Feng Lin for technical assistance in the flow cytometry analysis of rice tissues.

**Conflicts of Interest:** The authors declare that they have no competing interests.
