**5. Conclusions**

Answering question (i), the northern boundary of the Pannonian steppe-forest-steppe vegetation type is found in the study area, which is the valley of the Ipoly River. The vegetation type appears in the drier, acerbic sandy areas along the rivers, as well as new occurrence patches, and a special coenosystemic vegetation mixture is formed. (ii) In the study area, the nearby sand ridges provided an environmental background to learn about the boundaries of the Pannonian sand steppe and forest-steppe. The dominant species of the steppe patches was *Festuca rupicola*, which plays a similar coenosystemic role in the formation of the vegetation to *Festuca vaginata* in the central parts of the Carpathian Basin. Based on the data, the appearance of *Festuca vaginata* can also be expected due to global climatic changes [53–55]. (iii) Based on the Sentinel-2A data, we have seen that urban areas are separated from natural habitats but the individual vegetation types are not clearly separated.

**Author Contributions:** Conceptualization, I.J., F.S., and K.P.; Data curation, E.S.-F. and N.P.; Formal analysis, Z.Z.; Methodology, I.J., D.F., and P.C.; Software, D.S.; Supervision, K.P.; Writing—original draft, I.J.; Writing—review & editing, G.P. and K.P. All authors have read and agreed to the published version of the manuscript.

**Funding:** The work was funded by OTKA K-125423 and NKFIH-1159-6/2019.

**Institutional Review Board Statement:** Not applicable.

**Informed Consent Statement:** Not applicable.

**Data Availability Statement:** The data presented in this study are available on request from the corresponding author.

**Acknowledgments:** Our studies received financial support from the Higher Education Institutional Excellence Program (NKFIH-1159-6/2019) awarded by the Ministry for Innovation and Technology and by OTKA K-125423.

**Conflicts of Interest:** The authors declare no conflict of interest.
