3.2.2. Paratypes

A total of 27 specimens: Three adult males (MUSM 37355, MUSM 37357, UMMZ 246569) and an adult female (MUSM 37356) (Figures 3 and 4); from Los Amigos Biological Station, collected by R. von May and R. Santa Cruz on 26 November 2016; an adult female (UMMZ 246570) from Los Amigos Biological Station, collected by R. von May and R. Santa Cruz on 29 November 2016; three adults (sex unknown) (UMMZ 244945, MUSM 33247, MUSA 6302) from Los Amigos Biological Station, collected by R. von May and R. Santa Cruz on 14–19 May 2014; three specimens male/male/adult (MUSA 6974, MUSM 24219, MUSM 24251) from Los Amigos Biological Station, collected with the holotype by R. von May and R. Santa Cruz on 16 January 2012. Seven males: MUBI 17412, CORBIDI 17521, from San Pedro, Kosñipata Valley (13.04514S, 71.52922W, 1274 m), collected on 7 April 2018 by A. Catenazzi and M. I. Diaz; CORBIDI 17520, from San Pedro, Kosñipata Valley (13.04833S, 71.53178W, 1341 m), collected on 7 April 2018 by A. Catenazzi and M. I. Diaz; CORBIDI 17524 from San Pedro, Kosñipata Valley (13.03724S, 71.52798W, 1200 m), collected on 26 January 2009 by A. Catenazzi; MVZ:Herp:292685, MVZ:Herp:292686 from San Pedro, Kosñipata Valley (13.0541S, 71.54632W, 1370 m), collected on 24 January 2009 by A. Catenazzi; MVZ:Herp:292687, from San Pedro, Kosñipata Valley (13.05093S, 71.53711W, 1375 m), collected on 28 January 2009 by A. Catenazzi. One juvenile: MUBI 17413, from San Pedro, Kosñipata Valley (13.04514S, 71.52922W, 1274 m), collected on 7 April 2018 by A. Catenazzi and M. I. Diaz. Eight females: MUSM 30429, from San Pedro, Kosñipata Valley (13.04335S, 71.53027W, 1234 m), collected on 26 January 2009 by A. Catenazzi; CORBIDI 17523, from San Pedro, Kosñipata Valley (13.05044S, 71.53373W, 1342 m), collected on 26 January 2009 by A. Catenazzi; MUSM 27578, from San Pedro, Kosñipata Valley (13.05686S, 71.54081W, 1369 m), collected on 2 February 2008 by A. Catenazzi; MUSM 30426, MUSM 30427, MUSM 30428, MVZ:Herp:292684, from San Pedro, Kosñipata Valley (13.05193S, 71.5376W, 1376 m), collected on 24 January 2009 by A. Catenazzi; CORBIDI 17522, from San Pedro, Kosñipata Valley (13.05704S, 71.54778W, 1412 m), collected on 7 February 2008 by A. Catenazzi. Dorsal and ventral views of five paratypes (MUSA 6974, MUSM-37355, UMMZ 246569, MUSM 37356, UMMZ 246570), as well as the holotype, are presented in Figure 5. Three-dimensional reconstructions based on μCT data, from the skull and skeleton of two paratypes (MUSM 37355, MUSM 37356) are presented in Figure 6.

**Figure 3.** Photographs of live specimen of *Noblella losamigos* sp. n. ( **A**–**C**) Adult male paratype MUSM 37355 (SVL 9.5 mm). Photographs by Rudolf von May.

**Figure 4.** Photographs of live specimens of *Noblella losamigos* sp. n. (**A**,**B**) Adult male paratype MUSM 37357 (SVL 10.3 mm). (**C**,**D**) Adult female paratype MUSM 37356 (SVL 9.7 mm). Photographs by Roy Santa-Cruz.

**Figure 5.** Photographs of preserved specimens of *Noblella losamigos* sp. n. (**A**,**B**) Adult female holotype MUSA 6973 (SVL 10.7 mm); (**C**,**D**) adult male paratype MUSA 6974 (SVL 9.8 mm); (**E**,**F**) adult male paratype MUSM 37355 (SVL 9.5 mm); (**G**,**H**) adult male paratype UMMZ 246569 (SVL 9.6 mm); (**I**,**J**): adult female paratype MUSM 37356 (SVL 9.7 mm); and (**K**,**L**) adult female paratype UMMZ 246570 (SVL 11.3 mm). Photographs by Rudolf von May, Roy Santa-Cruz, and Courtney Whitcher.

**Figure 6.** Three-dimensional reconstructions based on μCT data, from the skull and skeleton of two specimens of *Noblella losamigos* sp. n. (**A**,**B**) Dorsal and ventral views of specimen MUSM 37355 (SVL 9.6 mm). (**C**,**D**) Dorsal and ventral views of specimen MUSM 37356 (SVL 9.7 mm).

## 3.2.3. Generic Placement

We assign this species to the genus *Noblella* based on its overall morphological resemblance with other species of *Noblella*, including the presence of T-shaped and pointed terminal phalanges (especially in toes). The genus *Noblella* Barbour, 1930 can be recognized by the following characters [3,7,15,40,41]: tympanic membrane differentiated (except in *N. duellmani* and *N. madreselva*); head narrower than body; cranial crests absent; dentigerous processes of vomers absent; finger I shorter than, or equal in length to, finger II; toe III shorter than toe V; tips of at least toes III–IV pointed; subarticular tubercles not protruding; conspicuous tarsal tubercle; dark inguinal spots present (except in *N. duellmani*); small body size (SVL < 22 mm). Additionally, our phylogenetic analysis indicates that the new species is closely related to other species of *Noblella* distributed in southern Peru (Figure 1), but also to several species of *Psychrophrynella*. However, given that *N. peruviana* and *P. bagrecito* (the type species of *Noblella* and *Psychrophrynella*, respectively) have not been included in phylogenetic analyses, the assignment remains tentative.
