3.2.8. Variation

The description of the coloration in life is based on notes taken in the field and photographs from multiple individuals (Figures 3, 4 and 8–10). The dorsum varies from ocher gold to light brown. Most individuals have an irregularly-shaped dark brown dorsal mark, and an interorbital bar. In some individuals, a light middorsal line extending from the middle of the body to the cloaca replaces the brown dorsal mark. Ventral coloration varies from transparent gray to black. Although all individuals have minute irregular cream flecks on venter, throat, thighs, hands, and feet, the sizes and concentration of these flecks are variable. Some males have black coloration extending from throat to mid venter.

**Figure 8.** Photographs of live specimens of *Noblella losamigos* sp. n. ( **A**) Dorsal view and (**B**) ventral view of MUBI 17413, adult male paratype (SVL 9.6 mm); ( **C**) dorsal view and ( **D**) ventral view of CORBIDI 17520, adult male paratype (SVL 11.1 mm); (**E**) dorsal view and (**F**) ventral view of CORBIDI 17521, adult male paratype (SVL 9.6 mm). Photographs by A. Catenazzi.

**Figure 9.** Photographs of live specimens of *Noblella losamigos* sp. n. (**A**) Dorsal view and (**B**) ventral view of MUSM 30427, adult female paratype (SVL 11.6 mm); (**C**) dorsal view and (**D**) ventral view of MVZ:Herp:292684, adult female paratype (SVL 13.2 mm); (**E**) dorsal view and (**F**) ventral view of CORBIDI 17522, adult female paratype (SVL 13.6 mm). Photographs by A. Catenazzi.

Several individuals have evident dark brown circular suprainguinal marks. The forelimb pattern varies from speckled to some individuals having a dark brown transverse bar forming a wristband. A dark facial mask is present in most individuals, though variable in shape and extent. In most individuals, the upper lips have between one and three cream flecks; if present, one of these spots is below the eyes. In some specimens, the facial mask merges with a dark lateral line that extends from the tip of the snout and almost reaches to the point of insertion of the thighs. If present, the lateral line is often broken into blotches by interruptions of the lighter flank color.

Our morphological data indicate that body size and shape of *N. losamigos* sp. n. vary with elevation. Body size in both males and females increases with increasing elevation (Figure 11a; OLS regression model, females R<sup>2</sup> = 0.81, df = 7, P < 0.001; OLS regression model, males R<sup>2</sup> = 0.28, df = 7, P = 0.045). In addition, the PCA projection of body size-corrected data indicates that body shapes of males and females vary with elevation (Figure 11b). Males and females found in the lowlands occupy a morphological space that is smaller and does not overlap with the highland population, whilst individuals found in the highlands occupy a larger and more variable morphological space.

**Figure 10.** Photographs of live specimens of *Noblella losamigos* sp. n. (**A**) Dorsal view and (**B**) ventral view of CORBIDI 17524, adult male paratype (SVL 9.8 mm); (**C**) dorsal view and (**D**) ventral view of MUSM 30426, adult female paratype (SVL 13.1 mm); (**E**) dorsal view and (**F**) ventral view of MUBI 17412, juvenile paratype (SVL 7.1 mm).

**Figure 11.** Multi-panel plot displaying variation of body size and body shape in *Noblella losamigos* sp. n. across elevations. (**a**) Body size in females and males tends to increase with increasing elevation. (**b**) Projection of Principal Component Analysis based on body-size-corrected data; body shape differs between females and males and varies with respect to elevation.
