*2.4. Environmental Data*

Four climatic variables retrieved from the WorldClim climate database [39] (mean annual temperature, annual precipitation, temperature seasonality, and precipitation seasonality) and four variables related to human pressures (agricultural land area, urban land area, land-cover diversity, and population density) were investigated to understand their relationship with functional richness and Rao's quadratic entropy. Landscape data (agricultural land area, urban land area, and land-cover diversity) were provided by the land-cover dataset CLC2000 [40], and human population density was obtained from HYDE Gridded Population version 3.1 [41]. The environmental data were reprojected and resampled to the same projection and resolution as the distribution data in QGIS version 2.10.0.
