*2.4. Genome-Wide Association Analysis*

We analyzed the SNPs in Axiom\_Arachis with a 58K array of the cultivated peanut using R software analysis tools. In the present study, the GAPIT package of R software— was used to conduct GWAS, and the enriched compressed mixed linear model (ECMLM) was selected for the analysis of association between SNPs and the phenotype data of interest [55]. The cutoff for significant association was a false discovery rate (FDR) adjusted *p*-value of less than 0.05. Candidate genes covering significantly associated SNPs were selected from the PEANUTBASE website tool (https://www.peanutbase.org) within a 150 kb region upstream or downstream of peak SNPs according to the linkage disequilibrium (LD) decay results.
