4.1. Observed Geomorphology
Results from this study show how expected longitudinal change in channel geomorphology can be altered independently or by combined bedrock channel constraints and human activity. For example, the mid-reaches of HCW were associated with increased bedrock channel constraints (
Figure 3), which according to Montgomery et al. [
52] indicates transport capacity in excess of sediment supply. Outside of the mid-reaches (i.e., Sites #1 and #5) where less than 3% presence of channel bedrock constraints were observed, signs of channel incision were apparent. Site #1 in the agricultural headwaters and Site #5 located the lower urban reaches of HCW (
Figure 3). Agricultural and urban land use explained nearly all the variance in average width to depth ratios, and maximum bank angle (0.896 ≤ R
2 ≥ 0.960; 0.020 ≤ p ≥ 0.052; n = 5). At Site #1 located in the agricultural headwaters (drainage area = 79.0 km
2), width to depth ratios increased at a rate of about 0.3 km
−1, and maximum bank angle decreased by about 0.5 degrees km
−1 as agricultural land use decreased from 100 to 56.9%, forested land use increased from 0 to 35.9%, and urban land use increased from 0 to 4.7% over 22.8 km stream distance (
Figure 4). With stream distance from agricultural Site #1 to sub-urban Site #3 located at the rural-urban interface of HCW, (drainage area = 116.2.0 km
2), width to depth ratios increased by a rate of about 0.01 m m
−1 km
−1, and maximum bank angle decreased by about 0.1 degrees km
−1 as agricultural land use decreased to 49.5%, forested land use decreased by 0.5%, and urban land use increased to 13.0% over 12.6 km stream distance (
Figure 4). Continuing downstream from sub-urban Site #3 to urban Site #5 located in the lower urban reaches (drainage area = 207.5 km
2), width to depth ratios decreased by about 0.2 km
−1, and maximum bank angle increased by about 0.2 degrees km
−1 as agricultural land use decreased to 38.4%, forested land use decreased to 33.1%, and urban land use increased to 26.5% over 17.6 km stream distance (
Figure 4). Bankfull depth increased by 0.13 m km
−1 at Site #1, 0.02 m km
−1 between Sites #1 and #3, and 0.15 m km
−1 between Sites #3 and #5. In combination, these results indicated increased channel incision at Site #1 in the agricultural headwaters where agricultural land use accounted for greater than 50% of total catchment area, and Site #5 the lower urban reaches where urban land use accounted for greater than 20% of total catchment area in HCW. Thus, Sites #1 and #5 were considered hot spots of channel incision due to increased rates of change in channel geomorphology metrics.
Previous studies have shown agricultural land use can result in channel incision through various mechanisms including, but not limited to, channelization, deforestation, and alterations to soil and channel hydraulics [
53,
54,
55,
56]. Simon et al. [
56] noted increased pore water pressure coupled to decreased sheer strength can lead to streambank erosion. Given the influence of soil hydraulic forces on streambank stability, agricultural areas are subject to increased bank failure where crop irrigation lowers water table levels in the vicinity of a stream. A study by Zaimes et al. [
54] showed incised stream channels were associated with increased bank mass wasting, streambank erosion and sediment load in Beak Creek, an agricultural land use dominated Midwestern stream located in central Iowa, USA. Streambank erosion rates varied from meandering row crop fields (387 mm year
−1), cattle and horse pastures (295 mm year
−1), and meandering forest buffered reaches (142 mm year
−1). Results indicated that forested riparian buffers would reduce streambank erosion by 72%. Streambank stabilizing root systems associated with riparian vegetation add roughness, reduce stream power, and enhance bank accretion [
56]. Results from the aforementioned studies are in agreement with results from the current work showing agricultural land use alterations to channel morphology of Midwestern streams.
Previous studies have also shown urban land use can cause channel incision via alterations to streamflow regimes [
24,
25,
57,
58,
59]. A literature review by Walsh et al. [
24] showed increased impervious surfaces associated with urban land use can cause increased volume and velocity of surface runoff and a flashy hydrologic streamflow response linked to increased bank wasting, channel incision, and scouring. For example, Jordan et al. [
60] showed urban land use alterations to flow caused a 9 to 61% increase of sediment yield due to channel incision and bank erosion in Berryessa Creek, California, USA. A study by Shields et al. [
59] quantified differences in channel incision, streamflow, water quality, and stream physical habitat between rural and urban catchments located in the Yazoo River basin, Mississippi, USA. Results showed urban land use was associated with decreased physical aquatic habitat, 6.4 times median rate of rise, 1.8 times channel depth, 3.5 times channel width, 2 to 3 times turbidity and suspended solids, 2 times fish species, and 4 times the amount of fish biomass per unit of effort [
59]. Results from the current work are a novel addition to previous studies considering the intensive sampling regimen (n = 561) that made possible the estimation of the rate of change in channel morphology and substrate composition across an agricultural-urban land use gradient.
4.2. Observed Streambed Substrate Frequency and Embeddedness
No channel morphology variable significantly explained the variance in substrate embeddedness of pools, riffles, and glides for each bank to bank sampling position (
Figure 5) highlighting the spatial complexity of streambed substrate composition in this study. Spatial complexity of substrate composition was, in part, shaped by the thalweg which meandered from bank to bank. The thalweg was generally associated with decreased substrate embeddedness due to increased stream velocity, and thus, increased sediment transport capacity. While much of the observed bank to bank variability in substrate embeddedness was attributed to thalweg position, results from the current work in combination with previous research in the region indicated longitudinal variability in substrate composition was attributed to the presence claypan soils and agricultural land use in the headwaters, increased bed slope and bedrock channel constraints in the mid-reaches, and the influence of urban land use associated with increased impervious surfaces in the lower reaches (
Figure 3).
Previous studies in HCW and elsewhere have shown claypan soils and agricultural land use are associated with increased surface runoff, soil erosion, and channel sediment supply [
14,
15,
29]. Lerch et al [
29] noted claypan soils corresponding to the Mexico-Leonard association consisting of an argillic soil horizon of smectitic mineralogy with clay content of 450–650 g kg
−1 formed at 10 to 50 cm depth are characterized by increased surface runoff. Willett et al. [
15] showed claypan soils were associated with increased bank sediment supply that accounted for 88% of total sediment supply. Streambank erosion was particularly high during winter months attributed to a combination of increased frequency and large magnitude flow events, freeze/thaw cycles, high antecedent moisture conditions, and lack of vegetation. In the current work, increased bank angle and substrate embeddedness were apparent during field sampling in the agricultural headwaters of HCW where claypan soils are present. Results from the current work are among the first to quantitatively characterize agricultural land use influence on substrate frequency and substrate embeddedness in the Central Claypan Region and point to a need to mitigate the influence claypan soils and agricultural land use on degradation of stream hydrogeomorphology in HCW. Thus, results from this study in combination with previous research have implications in other agricultural watersheds where near surface soil features (e.g., claypans, argillic horizons, or fragipans) have increased sediment supply.
While substrate embeddedness was greater in the agricultural headwaters, substrate embeddedness decreased as bed slope increased in the mid-reaches. Agricultural Site #1 was associated with 26 to 39% greater frequency of substrate smaller than GC, and 20 to 25% greater substrate embeddedness compared to Sites #2 to #4 in the mid-reaches of HCW (
Figure 6). Bed slope, width to depth ratio, and the frequency of substrate greater than GC were inversely related to substrate embeddedness (
Figure 4). Process-based understanding of the control of channel morphology on streamflow and sediment transport [
52] indicated the aforementioned general trends in the observed data were physically meaningful. Given that bed slope controls velocity of streamflow, stream capacity, and stream competence, it makes sense that Sites #2 to #4 located in the mid-reaches were generally associated with less substrate embeddedness and frequency of substrate less than GC in diameter compared to Sites #1 and #5. The mid-reaches of HCW were also associated with increased bedrock substrate and channel constraints (
Figure 3), which indicated transport capacity in excess of sediment supply [
52]. However, outside of the mid-reaches, increased substrate embeddedness was observed at agricultural Site # 1 and urban Site #5 (
Figure 3).
The greatest rate of change in substrate embeddedness with downstream distance was observed in riffles located in the lower urban reaches where bank height exceeded 7 m (
Figure 7). For example, percent embeddedness of riffle habitat decreased by at a rate of about 2% km
−1 as agricultural land use decreased from 100 to 56.9%, forested land use increased from 0 to 35.9%, and urban land use increased from 0 to 4.7% over 22.8 km stream distance at Site #1 (
Figure 7). However, percent embeddedness of riffles remained relatively constant (i.e., negligible rate of change) from agricultural Site #1 to sub-urban Site #3 located at the rural-urban interface where increased bedrock channel constraints were observed in HCW (
Figure 7). Continuing downstream from sub-urban Site #3 to urban Site #4, substrate embeddedness of riffles began to increase by about 1.4% km
−1 over approximately 8 km of stream distance. Further downstream, percent embeddedness of riffles increased rapidly by 5.3% km
−1 between urban Sites #4 and #5 (
Figure 7). Similar trends were observed in glides and pools as well (
Figure 7). Thus, these results showed increased rate of change in percent embeddedness linked to degraded physical habitat (riffles, glides and pools) at agricultural Site #1 and urban Site #5, with a disproportionate rate of increase of substrate embeddedness in riffle habitat of the lower urban reaches pointing to symptoms of urban stream syndrome in HCW.
The observed channel incision in the lower reaches of the current work was corroborated by other authors who also reported urban land use influence on channel morphology [
24,
60,
61]. Blann et al. [
53] discussed how increased channel incision disrupts hydrologically and ecologically important stream-floodplain-riparian connectivity. In urban catchments, channel incision is often attributed to increased impervious surfaces and engineered waterways that connect impervious surfaces directly to stream channels [
24,
25]. Increased impervious surfaces have been shown to increase the volume and velocity of surface runoff in HCW [
36] and elsewhere [
24,
25,
62]. Increased surface runoff commonly translates to alterations to flow regimes (i.e., flow frequency, magnitude, timing, duration and rate of change), sediment transport regimes, water quality, and aquatic ecosystem health [
24,
25]. Clearly, there is a need to mitigate land use alterations to channel morphology via channel stabilization efforts in HCW and similar catchments globally. However, Vietz et al. [
58] noted channel stabilization methods may not be sufficient to combat channel degradation in urbanized catchments. There is also a critical need for flow regime management efforts to reduce flow magnitude, frequency, and rate of change associated with alterations to channel morphology.
It was evident in the current work that simultaneously occurring agricultural and urban land uses exacerbated problems with substrate embeddedness in HCW particularly in the headwaters and lower urban reaches were bedrock constraints were less abundant. Previous studies also showed a general trend for suspended sediment and nutrients to decrease from Site #1 in the agricultural headwaters to Site #3 at the rural-urban interface of the watershed, and then, increase from Site #3 to urban Site #5 located near the watershed outlet in HCW [
14,
40,
41]. In fact, significant relationships were observed between substrate embeddedness values reported in the current work and concentrations of suspended sediment (R
2 = 0.798; p = 0.026), nitrite-N (R
2 = 0.975; p = 0.001), and ammonia-N (R
2 = 0.956; p = 0.002) reported in previous studies in HCW [
14,
41]. Notably, suspended sediment and total phosphorous yields were particularly high compared to other regions within the Mississippi River Basin. For example, observed suspended sediment yields exceeding 300 Mg km
−2 year
−1 were 54%, 80%, and 87% greater than sediment yields from the Ohio River, the Upper Mississippi River, and the Missouri River basins, respectively [
14]. Total phosphorous yields (0.979 kg ha
−1 year
−1) were also high for the region [
41]. In combination with results of LULC alterations to substrate embeddedness from the current work, results indicate increased sediment supply has degraded water quality, physical habitat, and aquatic refugia in HCW, especially in the agricultural headwaters and the lower urban reaches.
Collectively, previous studies showed streams of the Midwestern USA are commonly associated with increased sediment supply and degraded stream health [
13,
15,
57]. For example, Gellis et al. [
57] noted channel sources of sediment accounted for the majority (>50%) of bed sediment in 79% of 99 Midwestern watersheds sampled. Increased channel sediment supply and subsequent bed sedimentation can bury riffle habitat, reduce egg and fry survivorship, and lower prey densities [
63,
64]. Results from previous studies often focused on agricultural influence on stream bank erosion and sedimentation which was observed in the current work. However, it should be noted that urban land use may cause a greater influence on channel morphology and streambed sedimentation relative to agricultural land use. For example, substrate embeddedness was observed to decrease between Site #1 and #2 where agricultural land use accounted for 55% of total catchment area. Continuing downstream, substrate embeddedness began to increase between Sites #2 and #3 at the rural—urban interface of HCW which was associated with about 7 to 13% urban land use in this study. Thus, results from this study were in agreement with Paul and Meyer [
65] who noted urban land use can account for a small percentage of total catchment area while causing a disproportionate influence on water quality and stream health relative to other land uses.