Description of a New Species of the Pareas hamptoni Complex from Yunnan, China, with Confirmation of P. hamptoni Sensu Stricto in China (Squamata, Pareidae) †
Abstract
Simple Summary
Abstract
1. Introduction
2. Materials and Methods
2.1. Sampling
2.2. Morphometrics
2.3. Molecular Analyses
3. Results
3.1. Morphological Results
3.2. Molecular Results
3.3. Systematics
4. Discussion
5. Conclusions
1 | Dorsal head solid black........................................................................................................................2 |
Dorsal head yellowish brown to reddish brown.............................................................................3 | |
2 | Dorsal body almost solid black........................................................................................Pareas niger |
Dorsal body yellow to brownish red with black bars......................................Pareas yunnanensis | |
3 | Vertebral scales not enlarged..................................................................................Pareas xuelinensis |
Three rows of vertebral scales enlarged...............................................................Pareas formosensis | |
One row of vertebral scales enlarged................................................................................................4 | |
4 | Subcaudals more than 91....................................................................Pareas hamptoni sensu stricto |
Subcaudals less than 91.......................................................................................................................5 | |
5 | Ventrals less than 188................................................................................................Pareas geminatus |
Ventrals more than 189..................................................................Pareas guanyinshanensis sp. nov. |
Author Contributions
Funding
Institutional Review Board Statement
Informed Consent Statement
Data Availability Statement
Acknowledgments
Conflicts of Interest
References
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Species | Voucher | Locality | Cyt b | ND4 |
---|---|---|---|---|
Pareas guanyinshanensis sp. nov. | KIZ 2023038 | Yuanyang, Yunnan, China | PP215390 | PP215399 |
Pareas guanyinshanensis sp. nov. | KIZ 2023039 | Yuanyang, Yunnan, China | PP215389 | PP215398 |
Pareas guanyinshanensis sp. nov. | KIZ 2023040 | Yuanyang, Yunnan, China | PP215388 | PP215397 |
Pareas abros | ZMMU R-16393 | Song Thanh, Quang Nam, Vietnam | MZ712235 | MZ712262 |
Pareas andersonii | CAS 235359 | Natmataung, Chin, Myanmar | MT968772 | MW287040 |
Pareas atayal | NMNS 05594 | Beiheng, Taoyuan, Taiwan | KJ642124 | MW287041 |
Pareas baiseensis | ANU 000220008 | Baise, Guangxi, China | OQ054328 | OQ054329 |
Pareas berdmorei | AUP 01573 | Chiang Mai, Thailand | MZ712218 | MZ712244 |
Pareas boulengeri | GP2923 | Jiangkou, Guizhou, China | MK135090 | MK805355 |
Pareas carinatus | KIZ 011972 | Malaysia | MK135111 | MK805376 |
Pareas chinensis | CIB 010140 | Tianquan, Sichuan, China | JF827691 | JF827669 |
Pareas dulongjiangensis | KIZ R201607 | Gongshan, Yunnan, China | OQ718498 | — |
Pareas formosensis | NMNS 05637 | Nantou, Taiwan | MW287060 | MW287042 |
Pareas formosensis | H26-HAM01 | Guangdong, China | MW287061 | MW287043 |
Pareas formosensis | YBU 12015 | Hainan, China | MK135068 | MK805333 |
Pareas formosensis | GP4581 | Jingning, Zhejiang, China | MK135072 | MK805337 |
Pareas formosensis | YBU 12115 | Rongjiang, Guizhou, China | MK135075 | MK805340 |
Pareas formosensis | YBU 14508 | Guangxi, China | MK135076 | MK805341 |
Pareas formosensis | GP3696 | Yanshan, Jiangxi, China | MH046857 | MK805382 |
Pareas formosensis | ZMMU R-16333 | Kon Chu Rang, Gia Lai, Vietnam | MW287066 | MW287048 |
Pareas formosensis | ZMMU NAP-08868 | Song Thanh, Quang Nam, Vietnam | MW287063 | MW287045 |
Pareas formosensis | ZMMU R-16684 | Phia Oac, Cao Bang, Vietnam | MW287062 | MW287044 |
Pareas formosensis | ZMMU R-13709 | Bidoup–Nui Ba, Lam Dong, Vietnam | MW287064 | MW287046 |
Pareas formosensis | ZMMU R-14072 | Chu Yang Sin, Dak Lak, Vietnam | MW287065 | MW287047 |
Pareas formosensis | FMNH 255567 | Pu Mat, Nghe An, Vietnam | AY425806 | — |
Pareas geminatus | DL2019072910 | Jiangcheng, Yunnan, China | MW287067 | — |
Pareas geminatus | DL2019093001 | Jiangcheng, Yunnan, China | MW287071 | — |
Pareas geminatus | DL2019093002 | Jiangcheng, Yunnan, China | MW287072 | — |
Pareas geminatus | CIB 118021 | Jiangcheng, Yunnan, China | MW287068 | — |
Pareas geminatus | CIB 118022 | Jiangcheng, Yunnan, China | MW287069 | — |
Pareas geminatus | CIB 118023 | Jiangcheng, Yunnan, China | MW287070 | — |
Pareas geminatus | KIZ L2020020 | Jiangcheng, Yunnan, China | MW436707 | — |
Pareas geminatus | KIZ L2020024 | Jiangcheng, Yunnan, China | MW436708 | — |
Pareas hamptoni sensu stricto | YPX 18604 | Kachin, Myanmar | MK135078 | MK805343 |
Pareas hamptoni sensu stricto | YPX 18219 | Kachin, Myanmar | MK135077 | MK805342 |
Pareas hamptoni sensu stricto | CAS 221489 | Putao, Kachin, Myanmar | MW287077 | — |
Pareas hamptoni sensu stricto | ZMMU NAP-09087 | Bat Xat, Lao Cai, Vietnam | MW287078 | MW287054 |
Pareas hamptoni sensu stricto | ZMMU NAP-09088 | Bat Xat, Lao Cai, Vietnam | MW287079 | MW287053 |
Pareas hamptoni sensu stricto | ROM 38104 | Sa Pa, Lao Cai, Vietnam | KX694896 | — |
Pareas hamptoni sensu stricto | KIZ 20210517 | Malipo, Yunnan, China | PP215386 | PP215395 |
Pareas hamptoni sensu stricto | KIZ L2020018 | Malipo, Yunnan, China | PP215382 | PP215391 |
Pareas hamptoni sensu stricto | KIZ 201903001 | Hekou, Yunnan, China | PP215383 | PP215392 |
Pareas hamptoni sensu stricto | KIZ 20210518 | Hekou, Yunnan, China | PP215385 | PP215394 |
Pareas hamptoni sensu stricto | KIZ 20210519 | Hekou, Yunnan, China | PP215384 | PP215393 |
Pareas hamptoni sensu stricto | KIZ 2023046 | Jianshui, Yunnan, China | PP215387 | PP215396 |
Pareas iwasakii | NMNS 05655 | Ishigaki, Okinawa, Japan | KJ642160 | — |
Pareas kaduri | BNHS 3574 | Lohit, Arunachal Pradesh, India | MW026190 | — |
Pareas komaii | NMNS 05618 | Lijia, Taitung, Taiwan | KJ642185 | MW287056 |
Pareas kuznetsovorum | ZMMU R-16802 | Song Hinh, Phu Yen, Vietnam | MZ712232 | MZ712258 |
Pareas macularius | ZMMU R-16629 | Ban Mauk, Sagaing, Myanmar | MT968771 | MW287057 |
Pareas margaritophorus | ZMMU NAP-09759 | Suan Phueng, Ratchaburi, Thailand | MZ712217 | MZ712243 |
Pareas modestus | MZMU 1293 | Aizawl, Mizoram, India | MT968773 | — |
Pareas monticola | ZMMU R-16631 | Ban Mauk, Sagaing, Myanmar | MW438296 | MW438301 |
Pareas niger | KIZ 059339 | Kunming, Yunnan, China | MW436706 | — |
Pareas niger | GP1294 | Mengzi, Yunnan, China | MK135079 | MK805344 |
Pareas niger | YBU 14251 | Mengzi, Yunnan, China | MK135080 | MK805345 |
Pareas niger | YBU 14252 | Mengzi, Yunnan, China | MK135081 | MK805346 |
Pareas niger | YBU 14253 | Mengzi, Yunnan, China | MK135082 | MK805347 |
Pareas niger | YBU 14288 | Mengzi, Yunnan, China | MK135083 | MK805348 |
Pareas niger | YBU 15100 | Kaiyuan, Yunnan, China | MK135084 | MK805349 |
Pareas niger | YBU 15114 | Kaiyuan, Yunnan, China | MK135085 | MK805350 |
Pareas nigriceps | CHS656 | Tengchong, Yunnan, China | MK201455 | — |
Pareas nuchalis | FK 2626 | Belait, Brunei Darussalam, Brunei | MZ603794 | U49311 |
Pareas stanleyi | HM 2007-S001 | Guilin, Guangxi, China | JN230704 | JN230705 |
Pareas temporalis | ZMMU R-13656 | Cat Loc, Lam Dong, Vietnam | MZ712238 | MZ712265 |
Pareas tigerinus | KIZ 20210703 | Menghai, Yunnan, China | OP752143 | — |
Pareas victorianus | CAS 235254 | Natmataung, Chin, Myanmar | MW438300 | MW438302 |
Pareas vindumi | CAS 248147 | Lukpwi, Kachin, Myanmar | MW287080 | MW287059 |
Pareas xuelinensis | KIZ XL1 | Lancang, Yunnan, China | MW436709 | — |
Pareas xuelinensis | KIZ XL2 | Lancang, Yunnan, China | MW436710 | — |
Pareas yunnanensis | KIZ 2022033 | Dali, Yunnan, China | OP752146 | — |
Pareas yunnanensis | KIZ 2022034 | Dali, Yunnan, China | OP752147 | — |
Pareas yunnanensis | KIZ 2022035 | Dali, Yunnan, China | OP752148 | — |
Pareas yunnanensis | KIZ 2022036 | Dali, Yunnan, China | OP752149 | — |
Aplopeltura boa | LSUHC 7248 | Sepilok, Sabah, Malaysia | KC916746 | U49312 |
KIZ 2023038 Holotype ♀ | KIZ 2023039 Paratype ♀ | KIZ 2023040 Paratype ♀ | |
---|---|---|---|
SVL | 488 | 482 | 540 |
TL | 146 | 126 | 152 |
PrFBO | Yes | Yes | Yes |
PreO | 1/1 | 1/1 | 1/1 |
PosO | 1/1 | Fused/1 | 1/Fused |
SubO | 1/1 | Fused/1 | 1/Fused |
SPOF | No/No | Yes/No | No/Yes |
ATem | 2/2 | 2/2 | 2/2 |
PTem | 3/3 | 4/4 | 2/2 |
SupL | 7/7 | 8/8 | 7/7 |
InfL | 7/8 | 7/6 | 7/7 |
LoBO | No | No | No |
Vs | 190 | 192 | 189 |
Prec | Undivided | Undivided | Undivided |
Sc | 89 | 72 | 80 |
DS | 15-15-15 | 15-15-15 | 15-15-15 |
NED | 1 | 1 | 1 |
NKD | 0-5-5 | 0-5-7 | 0-5-5 |
Max | 5/4 | 5/5 | 5/5 |
KIZ 20210517 ♂ | KIZ L2020018 Juvenile | KIZ 201903001 ♀ | KIZ 20210518 Juvenile | KIZ 20210519 ♂ | KIZ 2023046 ♂ | |
---|---|---|---|---|---|---|
SVL | 458 | 276 | 360 | 306 | 459 | 532 |
TL | 157 | 91 | 114 | 97 | 151 | Incomplete |
PrFBO | Yes | Yes | Yes | Yes | Yes | Yes |
PreO | 1/1 | 1/1 | 1/1 | 1/1 | 1/1 | 1/1 |
PosO | Fused/Fused | Fused/Fused | Fused/Fused | 1/1 | 1/1 | 1/1 |
SubO | Fused/Fused | Fused/Fused | Fused/Fused | 1/1 | 1/1 | 1/1 |
SPOF | Yes/yes | Yes/yes | Yes/yes | No/No | No/No | No/No |
ATem | 2/2 | 2/2 | 1/1 | 2/2 | 3/3 | 2/2 |
PTem | 3/2 | 3/3 | 3/2 | 2/3 | 3/3 | 3/3 |
SupL | 7/8 | 7/7 | 7/7 | 7/7 | 8/7 | 7/7 |
InfL | 6/6 | 7/7 | 7/7 | 7/7 | 8/8 | 7/7 |
LoBO | No | No | No | No | No | No |
Vs | 188 | 188 | 187 | 193 | 191 | 186 |
Prec | Undivided | Undivided | Undivided | Undivided | Undivided | Undivided |
Sc | 92 | 97 | 94 | 95 | 91 | Incomplete |
DS | 15-15-15 | 15-15-15 | 15-15-15 | 15-15-15 | 15-15-15 | 15-15-15 |
NED | 1 | 1 | 1 | 1 | 1 | 1 |
NKD | 0-3-5 | 0-3-3 | 0-3-5 | 0-3-3 | 0-3-5 | 0-5-7 |
Max | 4/5 | 5/5 | 5/5 | 4/4 | 5/5 | 4/5 |
(1) | (2) | (3) | (4) | (5) | (6) | (7) | (8) | (9) | (10) | (11) | (12) | (13) | (14) | (15) | (16) | (17) | (18) | (19) | (20) | (21) | (22) | (23) | (24) | (25) | (26) | (27) | (28) | (29) | (30) | |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
(1) | ||||||||||||||||||||||||||||||
(2) | 23.5 | |||||||||||||||||||||||||||||
(3) | 21.5 | 23.5 | ||||||||||||||||||||||||||||
(4) | 13.6 | 22.8 | 20.3 | |||||||||||||||||||||||||||
(5) | 19.2 | 23.7 | 20.6 | 20.6 | ||||||||||||||||||||||||||
(6) | 23.1 | 21.0 | 23.8 | 23.5 | 22.5 | |||||||||||||||||||||||||
(7) | 17.2 | 23.2 | 19.7 | 18.4 | 14.2 | 23.6 | ||||||||||||||||||||||||
(8) | 23.1 | 21.8 | 22.9 | 23.2 | 22.4 | 13.9 | 22.6 | |||||||||||||||||||||||
(9) | 18.0 | 23.8 | 19.2 | 18.8 | 14.6 | 24.9 | 9.0 | 22.8 | ||||||||||||||||||||||
(10) | 12.8 | 24.0 | 19.6 | 13.9 | 19.6 | 24.2 | 18.0 | 23.7 | 17.8 | |||||||||||||||||||||
(11) | 8.0 | 23.9 | 21.6 | 15.0 | 19.2 | 24.3 | 17.1 | 23.9 | 17.8 | 13.4 | ||||||||||||||||||||
(12) | 7.5 | 22.9 | 22.2 | 14.1 | 19.0 | 22.9 | 17.4 | 23.2 | 19.0 | 12.8 | 9.5 | |||||||||||||||||||
(13) | 4.2 | 23.5 | 21.7 | 14.0 | 18.9 | 23.3 | 17.2 | 23.5 | 18.3 | 12.6 | 8.0 | 7.4 | ||||||||||||||||||
(14) | 13.1 | 23.4 | 20.5 | 7.0 | 19.3 | 24.4 | 17.4 | 24.3 | 18.3 | 14.0 | 14.5 | 14.5 | 13.7 | |||||||||||||||||
(15) | 12.3 | 24.5 | 20.2 | 14.7 | 19.8 | 24.5 | 19.4 | 22.2 | 18.8 | 9.1 | 13.4 | 13.4 | 12.7 | 14.8 | ||||||||||||||||
(16) | 14.3 | 23.2 | 19.5 | 8.6 | 19.5 | 23.9 | 18.0 | 23.9 | 18.2 | 13.7 | 14.7 | 14.9 | 14.4 | 8.1 | 15.5 | |||||||||||||||
(17) | 22.7 | 21.2 | 23.8 | 23.2 | 23.3 | 13.2 | 22.6 | 13.2 | 23.0 | 24.2 | 23.3 | 23.1 | 23.2 | 24.4 | 22.9 | 24.3 | ||||||||||||||
(18) | 18.7 | 23.9 | 14.5 | 18.9 | 18.5 | 22.7 | 18.4 | 21.7 | 17.6 | 18.3 | 19.8 | 20.3 | 18.9 | 19.5 | 19.6 | 19.2 | 22.6 | |||||||||||||
(19) | 19.9 | 24.7 | 16.2 | 18.9 | 20.0 | 24.2 | 19.2 | 22.8 | 18.7 | 17.9 | 19.6 | 20.5 | 19.4 | 19.0 | 20.1 | 18.4 | 23.1 | 14.9 | ||||||||||||
(20) | 20.4 | 23.5 | 12.0 | 18.4 | 19.1 | 24.2 | 19.2 | 24.1 | 18.8 | 18.6 | 20.3 | 20.1 | 19.7 | 19.8 | 19.2 | 17.9 | 24.4 | 12.0 | 14.8 | |||||||||||
(21) | 18.6 | 22.7 | 18.7 | 17.7 | 20.3 | 21.7 | 18.3 | 22.9 | 18.5 | 17.6 | 19.5 | 19.5 | 18.6 | 17.7 | 18.7 | 18.1 | 21.9 | 17.1 | 18.9 | 18.7 | ||||||||||
(22) | 5.7 | 22.6 | 20.6 | 14.1 | 19.0 | 23.5 | 17.6 | 23.3 | 17.9 | 12.2 | 8.3 | 6.9 | 6.0 | 13.6 | 12.1 | 14.6 | 22.7 | 18.6 | 19.9 | 19.0 | 18.9 | |||||||||
(23) | 12.4 | 23.6 | 18.8 | 15.7 | 16.9 | 22.9 | 16.9 | 22.9 | 16.2 | 10.3 | 13.3 | 13.5 | 12.6 | 16.1 | 9.7 | 16.2 | 23.9 | 19.3 | 17.8 | 16.4 | 18.8 | 12.6 | ||||||||
(24) | 25.0 | 21.1 | 24.3 | 23.4 | 23.4 | 21.2 | 24.3 | 21.2 | 24.1 | 24.6 | 24.6 | 25.1 | 24.8 | 24.3 | 25.0 | 23.5 | 20.4 | 23.3 | 24.6 | 24.5 | 22.5 | 25.2 | 23.8 | |||||||
(25) | 19.2 | 25.7 | 20.4 | 19.2 | 16.3 | 25.2 | 15.7 | 24.7 | 15.4 | 19.2 | 19.5 | 19.5 | 18.7 | 18.9 | 20.3 | 17.3 | 24.9 | 18.6 | 19.5 | 19.4 | 18.8 | 19.6 | 19.0 | 24.0 | ||||||
(26) | 23.8 | 13.3 | 23.8 | 23.6 | 23.3 | 20.6 | 22.5 | 20.1 | 21.9 | 25.1 | 24.6 | 23.9 | 23.7 | 23.6 | 24.8 | 24.2 | 20.1 | 23.4 | 24.8 | 23.6 | 23.0 | 23.6 | 24.1 | 20.2 | 24.1 | |||||
(27) | 12.1 | 23.0 | 19.3 | 14.3 | 19.7 | 24.1 | 19.0 | 23.3 | 18.7 | 10.3 | 12.9 | 12.4 | 11.8 | 14.2 | 10.7 | 14.0 | 24.3 | 18.6 | 19.5 | 18.2 | 18.7 | 11.4 | 4.3 | 25.0 | 19.3 | 24.8 | ||||
(28) | 19.1 | 24.3 | 20.6 | 19.5 | 19.2 | 22.8 | 19.1 | 22.9 | 17.5 | 18.0 | 18.1 | 18.6 | 18.6 | 19.9 | 18.4 | 19.3 | 22.9 | 19.0 | 20.6 | 19.3 | 14.3 | 17.8 | 19.1 | 24.7 | 19.0 | 24.5 | 17.9 | |||
(29) | 11.0 | 24.4 | 20.8 | 14.7 | 19.5 | 24.4 | 18.4 | 24.2 | 17.7 | 12.6 | 12.5 | 12.4 | 11.5 | 14.5 | 12.3 | 15.0 | 23.8 | 19.0 | 19.9 | 19.9 | 17.9 | 11.0 | 12.3 | 24.7 | 19.4 | 25.3 | 11.9 | 17.8 | ||
(30) | 8.3 | 23.1 | 21.3 | 13.8 | 20.5 | 24.7 | 16.9 | 24.3 | 18.7 | 13.3 | 8.9 | 6.0 | 8.3 | 13.9 | 13.2 | 14.8 | 24.6 | 19.7 | 20.0 | 20.2 | 19.6 | 7.3 | 12.5 | 25.9 | 19.5 | 24.7 | 12.3 | 18.8 | 12.6 | |
(31) | 7.1 | 23.2 | 22.1 | 14.4 | 19.1 | 24.1 | 16.7 | 23.5 | 18.2 | 12.9 | 8.6 | 4.1 | 6.2 | 13.9 | 12.4 | 14.7 | 23.7 | 20.1 | 20.1 | 20.6 | 19.6 | 6.4 | 12.8 | 24.9 | 19.5 | 23.9 | 11.7 | 18.7 | 11.5 | 6.2 |
(1) | (2) | (3) | (4) | (5) | (6) | (7) | (8) | (9) | (10) | (11) | (12) | (13) | (14) | (15) | (16) | (17) | (18) | (19) | (20) | (21) | |
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
(1) | |||||||||||||||||||||
(2) | 18.6 | ||||||||||||||||||||
(3) | 19.0 | 20.4 | |||||||||||||||||||
(4) | 16.7 | 22.3 | 19.1 | ||||||||||||||||||
(5) | 18.1 | 21.8 | 18.0 | 20.2 | |||||||||||||||||
(6) | 21.0 | 19.5 | 21.5 | 22.0 | 25.5 | ||||||||||||||||
(7) | 18.7 | 21.1 | 17.8 | 18.4 | 12.1 | 22.8 | |||||||||||||||
(8) | 21.5 | 18.9 | 23.0 | 21.1 | 22.4 | 14.3 | 21.0 | ||||||||||||||
(9) | 18.3 | 21.1 | 18.3 | 20.1 | 12.2 | 22.8 | 10.1 | 22.6 | |||||||||||||
(10) | 9.1 | 18.8 | 19.6 | 16.2 | 17.1 | 20.7 | 19.3 | 21.3 | 17.5 | ||||||||||||
(11) | 5.0 | 19.4 | 19.0 | 16.4 | 17.5 | 21.2 | 17.9 | 20.9 | 16.5 | 8.7 | |||||||||||
(12) | 17.8 | 21.9 | 19.7 | 7.5 | 20.8 | 22.3 | 19.0 | 21.6 | 19.2 | 16.2 | 17.5 | ||||||||||
(13) | 19.1 | 19.9 | 21.3 | 21.1 | 22.8 | 14.1 | 19.8 | 15.0 | 21.7 | 19.7 | 18.9 | 21.3 | |||||||||
(14) | 20.3 | 22.5 | 11.8 | 19.8 | 18.6 | 22.9 | 19.5 | 23.2 | 20.2 | 20.9 | 20.4 | 20.8 | 23.7 | ||||||||
(15) | 19.9 | 20.8 | 14.0 | 19.6 | 18.9 | 21.3 | 18.6 | 21.0 | 20.8 | 19.6 | 20.1 | 21.0 | 20.2 | 14.7 | |||||||
(16) | 18.7 | 21.7 | 19.6 | 18.9 | 19.3 | 21.6 | 18.2 | 20.7 | 18.4 | 20.0 | 18.5 | 20.5 | 19.5 | 22.0 | 21.4 | ||||||
(17) | 9.5 | 19.8 | 18.8 | 16.0 | 18.0 | 22.2 | 17.6 | 21.4 | 17.1 | 8.8 | 9.0 | 16.2 | 21.3 | 20.4 | 18.6 | 19.7 | |||||
(18) | 21.3 | 17.4 | 19.9 | 23.4 | 22.8 | 18.2 | 22.3 | 18.8 | 22.5 | 20.3 | 21.1 | 23.7 | 19.5 | 23.4 | 21.1 | 21.3 | 21.4 | ||||
(19) | 20.3 | 21.6 | 18.0 | 19.5 | 16.2 | 23.9 | 14.8 | 22.5 | 16.6 | 19.3 | 20.4 | 18.6 | 22.4 | 18.0 | 19.5 | 19.2 | 19.1 | 23.0 | |||
(20) | 19.0 | 9.4 | 20.0 | 19.9 | 20.8 | 19.3 | 20.2 | 19.2 | 18.7 | 18.9 | 19.4 | 19.6 | 18.9 | 21.9 | 19.6 | 19.5 | 18.0 | 17.4 | 20.1 | ||
(21) | 17.6 | 20.4 | 18.5 | 18.4 | 18.4 | 22.9 | 18.3 | 20.8 | 18.6 | 18.0 | 17.4 | 19.0 | 21.7 | 20.5 | 20.2 | 12.8 | 18.6 | 23.5 | 19.5 | 18.3 | |
(22) | 12.5 | 19.0 | 16.3 | 13.9 | 14.8 | 21.1 | 15.6 | 20.4 | 15.7 | 12.8 | 13.0 | 14.2 | 19.2 | 18.0 | 18.6 | 17.4 | 13.6 | 20.1 | 16.5 | 18.5 | 15.6 |
Pareas guanyinshanensis sp. nov. | Pareas hamptoni Sensu Stricto | |
---|---|---|
SVL (adult) | 482–540 | 360–532 |
TL (adult) | 126–152 | 114–157 |
TL/SVL | 0.26–0.30 | 0.32–0.37 |
PrFBO | Yes | Yes |
PreO | 1 | 1 |
PosO | Fused or 1 | Fused or 1 |
SubO | Fused or 1 | Fused or 1 |
SPOF | Yes or No | Yes or No |
ATem | 2 | 1–3 |
PTem | 2–4 | 2–3 |
SupL | 7–8 | 7–8 |
InfL | 6–8 | 6–9 |
LoBO | No | No |
Vs | 189–192 | 185–195 |
Prec | Undivided | Undivided |
Sc | 72–89 | 91–99 |
DS | 15-15-15 | 15-15-15 |
NED | 1 | 1 |
NKD | 0–5 | 0–9 |
Max | 4–5 | 4–5 |
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© 2024 by the authors. Licensee MDPI, Basel, Switzerland. This article is an open access article distributed under the terms and conditions of the Creative Commons Attribution (CC BY) license (https://creativecommons.org/licenses/by/4.0/).
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Liu, S.; Mo, M.; Li, M.; Li, B.; Luo, X.; Rao, D.; Li, S. Description of a New Species of the Pareas hamptoni Complex from Yunnan, China, with Confirmation of P. hamptoni Sensu Stricto in China (Squamata, Pareidae). Animals 2024, 14, 421. https://doi.org/10.3390/ani14030421
Liu S, Mo M, Li M, Li B, Luo X, Rao D, Li S. Description of a New Species of the Pareas hamptoni Complex from Yunnan, China, with Confirmation of P. hamptoni Sensu Stricto in China (Squamata, Pareidae). Animals. 2024; 14(3):421. https://doi.org/10.3390/ani14030421
Chicago/Turabian StyleLiu, Shuo, Mingzhong Mo, Mei Li, Biao Li, Xiong Luo, Dingqi Rao, and Song Li. 2024. "Description of a New Species of the Pareas hamptoni Complex from Yunnan, China, with Confirmation of P. hamptoni Sensu Stricto in China (Squamata, Pareidae)" Animals 14, no. 3: 421. https://doi.org/10.3390/ani14030421
APA StyleLiu, S., Mo, M., Li, M., Li, B., Luo, X., Rao, D., & Li, S. (2024). Description of a New Species of the Pareas hamptoni Complex from Yunnan, China, with Confirmation of P. hamptoni Sensu Stricto in China (Squamata, Pareidae). Animals, 14(3), 421. https://doi.org/10.3390/ani14030421