Search Results (48)

Climate change threatens the stability of temperate grassland ecosystems in Inner Mongolia, a core part of the Eurasian Steppe, by driving widespread shifts in plant species distributions. Agropyron cristatum (L.) Gaertn., a dominant native perennial herb in Inner Mongolian steppes, is ecologically vital for degraded grassland restoration and forage supply, but its response to future climate change is unclear. Here, we used an optimized MaxEnt model to assess its potential distribution under current and future climate scenarios. We processed 228 initial occurrence records into 112 valid points, selected 11 non-collinear environmental variables, optimized model parameters with the R package ENMeval, and projected distributions for the 2050s and 2100s under CMIP6 SSP2-4.5 and SSP5-8.5 scenarios, while quantifying habitat fragmentation with landscape metrics. We found that annual mean temperature and annual precipitation dominate A. cristatum distribution (total contribution ~87%), with current highly suitable habitats concentrated in central-eastern Inner Mongolia. Future scenarios show stable core suitable habitats with northward and westward shifts, habitat fragmentation will slightly increase. Our findings clarify the climate response of A. cristatum and support its conservation and adaptive grassland management. Full article

Artemisia frigida (subshrub) communities, which are indicators of grassland degradation, are widespread in overgrazed Eurasian steppes. After 4–6 years of enclosure, the community can recover to an Agropyron cristatum-dominated grass community. Understanding the competitive mechanisms between these two key species provides critical insights for the management of semi-arid steppes, where vegetation dynamics are primarily driven by soil moisture. Nevertheless, how soil moisture distribution mediates above- and belowground competition between A. cristatum and A. frigida remains unclear. To address this, we conducted a pot experiment that simulated natural vertical soil moisture heterogeneity with four soil moisture regimes in two soil layers (0–30 cm and 30–60 cm): uniformly dry (D–D), wet upper/dry lower (W–D), dry upper/wet lower (D–W), and uniformly wet (W–W), using both monoculture and mixed planting methods. Key results showed that (1) A. cristatum was more sensitive to soil moisture regimes than A. frigida. Its above- and belowground biomass were significantly higher under moist treatments (W–W, W–D, D–W) than under drought (D–D), whereas the biomass of A. frigida did not differ significantly among water treatments. (2) Compared with monoculture, mixed planting significantly increased the root–shoot ratio of A. frigida but did not affect that of A. cristatum. (3) Competitive ability differed between aboveground and belowground parts: competitive indices (aggressivity and relative competition intensity) revealed that A. cristatum exhibited stronger aboveground competitiveness under moist treatments, while A. frigida dominated aboveground under drought conditions. However, A. frigida consistently exhibited greater belowground competitive ability than A. cristatum across all water treatments in the mixture. These results emphasize that assessments of grass–shrub competition based solely on aboveground indicators may underestimate the competitive advantage of shrubs. Therefore, integrating belowground competitive processes is essential for accurately predicting grass–shrub competition and succession in semi-arid steppes. Full article

Grassland productivity faces heightened uncertainty under nonlinear climatic forcing. This study characterizes the spatial heterogeneity of nonlinear variations and nonstationary climate sensitivities across the Eurasian Steppe Region (EASR) to provide a scientific basis for its adaptive management. Using the aboveground net primary productivity (ANPP) and climate datasets (1982–2015), we employed piecewise linear regression, LOWESS, and sliding window partial correlation analysis to identify temporal turning points and dynamic climate–productivity relationships. We identified distinct turning points in 1994 and 2008, revealing a phased “Increasing–Decreasing–Increasing” trajectory. A key novelty is the mapping of eight phased trajectory patterns, illustrating significant spatial heterogeneity in productivity trends. Furthermore, we demonstrate temporally reversed climate sensitivities. Notably, the sensitivity of ANPP to temperature shifted from positive to negative as warming-induced water stress intensified. While precipitation remains the dominant driver (68% of the region), its influence is nonstationary and state-dependent. In the Qinghai–Tibet Plateau, the limiting factor transitioned from thermal to water availability. Overall, productivity in the EASR appears to undergo phased reorganization under shifting climatic baselines. Our findings suggest that future ecosystem models should incorporate time-varying sensitivity parameters to account for nonlinear dynamics and potential trend reversals in grassland ecosystems. Full article

Heat waves (HWs), characterized by periods of unusually high temperature, would inevitably affect the soil microenvironment and then soil respiration (Rs), which is considered to be the most active part of the global carbon cycle. An in situ spring and summer HWs simulation experiment combined with a locally common human disturbance (mowing) was conducted to separate Rs into autotrophic respiration (Ra) and heterotrophic respiration (Rh) on a natural Eurasian meadow steppe in 2018 and 2019. HWs significantly affected grassland Rs, Rh and Ra (p < 0.01) and also interacted with mowing, but the effect of spring HW and summer HW were different. During the summer HWs, daily Rs of the non-mowed plots increased by 1.07 μmol m⁻² s⁻¹ (11.71%) and increased in the mowed plots by 2.15 μmol m⁻² s⁻¹ (23.81%). During the spring HWs, daily Rs of the non-mowed plots decreased by 0.13 μmol m⁻² s⁻¹ (2.36%) and decreased by 0.52 μmol m⁻² s⁻¹ (9.02%) in the mowed plots. Rs, Rh and Ra were inhibited by spring HWs, but promoted by summer HWs. Our results indicated that the occurring time change in HWs would cause widely divergent influences on the ecosystem, and mowing would decrease the anti-interference ability of the ecosystem, which acted as an enhancement on both the positive and negative effects of HWs. These findings have important implications for accurate model prediction and carbon budget assessment. Full article

This article reexamines the gold necklace excavated from the Sui-dynasty tomb of Li Jingxun (李静训, 600–608 CE), shifting attention from stylistic attribution to ritual function and funerary context. While previous studies have emphasized Persian, Byzantine, or Indian influences, this study situates the necklace more plausibly within the Iranian–steppe cultural sphere and the Turkic–Sogdian exchange networks active along the Silk Roads in the late sixth and early seventh centuries. Through analysis of its segmented structure, polyhedral gold beads, pearl rondelle, nicolo intaglio clasp, and gemstone arrangement, the article identifies close technical and visual parallels in Central Asia and the wider Iranian world. The necklace is interpreted as an apotropaic object likely worn in life and placed in the tomb to extend its protective and guiding functions after death. Attention to bodily use, clasp orientation, and associated grave goods—especially a stemmed cup with Eurasian ritual associations—clarifies how the necklace operated within a Buddhist burial setting timed to Lichun 立春 (Beginning of Spring). Situating the object within the Li family’s Xianbei 鲜卑 background and documented connections with Sogdian communities, this study demonstrates how foreign ornaments were actively understood and integrated into Sui aristocratic funerary practice, rather than adopted as passive luxuries. Full article

Hulun Lake, the largest freshwater lake in the Eurasian steppe, is a critically climate-sensitive water body facing severe ecological threats. This systematic review synthesizes multidisciplinary evidence from 1961 to 2025 to examine the characteristics and drivers of its water environment and quality evolution. The findings reveal that the primary driver of the lake’s hydrological degradation shifted from natural climate variability to anthropogenic land-use change around 1998. While ecological water diversion has partially alleviated water scarcity, it also introduces a significant external nutrient load, creating a paradox where increased water volume coincides with aggravated eutrophication. Furthermore, overgrazing in the catchment not only enhances conventional runoff pollution but also facilitates a unique “tumbleweed-mediated cross-media pollution” pathway. This review concludes that the restoration of Hulun Lake necessitates a shift from singular water quantity regulation to an integrated management strategy that concurrently addresses water quantity, quality, and aquatic ecosystem health. The insights gained are crucial for informing the sustainable management of Hulun Lake and other inland lakes in cold, arid regions under global change. Full article

In order to determine the impact of urbanization on raptors in the semi-desert conditions of southwestern Kazakhstan, an analysis of the spatio-temporal distribution of raptors is presented for the first time based on the results of surveys of the avifauna of Kyzylorda. Eight species of raptors were recorded: field Hen Harrier (Circus cyaneus), Marsh Harrier (C. aeroginosus), Eurasian Sparrowhawk (Accipiter nisus), Long-Legged Buzzard (Buteo rufinus), Eurasian Buzzard (B. buteo), Steppe Eagle (Aquila nipalensis), Eurasian Hobby (Falco subbuteo), and Common Kestrel (F. tinnunculus). The probability of raptors being present was negatively associated with dense urban low-rise buildings with limited greenery in the bay and the new part of the city. At the same time, the dense urban development with little or no greenery in the old central part of the city provides adequate habitat (including a foraging base and nesting sites) for the Common Kestrel. Raptor presence was positively associated with the Syrdarya River floodplain and wasteland with small groups of trees and/or shrubs. The landfill site located on the north-eastern edge of the city serves as a feeding ground for the Long-Legged and Eurasian Buzzards, while the airport area is inhabited by the Eurasian Buzzard, Steppe Eagle, and Common Kestrel. Based on this study, we would recommend that enterprises (e.g., grain storage facilities, airports) and local executive bodies who are interested in the conservation of raptors and regulating the population of the pigeons around their territories should retain or plant more native vegetation and shrubs and preserve areas with green spaces. Full article

This paper shows the results of a study on the ecological structure and spatial distribution of communities of the protected steppe ground beetle Carabus hungaricus Fabricius, 1792 (Coleoptera: Carabidae), in Bulgaria. It also points some phenological data and details about the species activity. In the period May 2021–December 2023, we explored 42 sample plots in xerophytic landscapes in central–western Bulgaria, representing the southernmost limits of the steppe fauna in Eurasia. With 252 pitfall traps, 69,903 effective trap-days were realised, and 15,333 carabid individuals from 184 species were identified. Carabus hungaricus was established in seven sites (frequency of occurrence, F = 16.7%), with a total of 198 specimens (1.3% of all carabids and almost 4% of the total dynamic density). In these seven sites, we found 57 species, mostly open habitat dwellers. The tribe Harpalini had 33% of the species. The most species-rich genera were Harpalus (13 species), Amara (8) and Carabus (6). Calathus distinguendus was superdominant with more than 71% of all carabid specimens. Euconstant species, collected in all seven sites (F = 100%), were Carabus coriaceus and Calathus distinguendus. The two classes of life forms had almost equal proportions, 28 (49.1%) Zoophagous and 29 (50.9%) Mixophytophagous species, which is extraordinary for Bulgaria, but typical for the Eurasian steppe zone. Macropterous beetles comprised 56% of the species and brachypterous were 25%, which is comparable only with typical montane habitats. Concerning humidity preferences, mesoxerophilous carabids predominated (25 species, 44%). In relation to its ground beetle fauna, the studied complex of habitats is quite remarkable for Bulgaria and appears to be stable and characteristic for the steppe biome. Full article

As a vital part of the Eurasian temperate grassland, the Chinese temperate grassland is primarily distributed in the Inner Mongolia Plateau. This paper focuses on mapping temperate grassland dynamics from the 1980s to the 2010s in Inner Mongolia, which was divided into temperate meadow steppe (TMS), temperate typical steppe (TTS), temperate desert steppe (TDS), temperate steppe desert (TSD) and temperate desert (TD). Multi-source features, including multispectral reflectance, vegetation growth, topography, water bodies, meteorological data, and soil characteristics, were selected based on their distinct physical properties and remote sensing variations. Then, we applied deep neural network (DNN) models to classify them, achieving an accuracy of 79.4% in the 1980s and 81.1% in the 2000s. Additionally, validation in the 2010s through field reconnaissance demonstrated an accuracy of 72.7%, which was acceptable, confirming that DNN is an effective method for classifying temperate grasslands. The results revealed that TTS had the highest proportion in the study area (39%), while TMS and TSD had the lowest (8.2% and 8.1%, respectively). Grassland types have the distribution law of aggregation; according to statistics, 61.1% of the grassland area remained unchanged, and the transition zone between adjacent grassland classes was highly easy to change. The area variation mainly came from TTS, TDS, and TSD, but not TD. The mutual transformation of different grassland types occurred mainly in adjacent areas between them. This study demonstrates the potential of DNN for long-term grassland mapping and provides the most comprehensive classification maps of Inner Mongolia grasslands to date, which are invaluable for grassland research and conservation efforts in the area. Full article

Unmanned aerial vehicles (UAVs) are rapidly becoming a popular tool for digital soil mapping at a large-scale. However, their applicability in areas with homogeneous vegetation (i.e., not bare soil) has not been fully investigated. In this study, we aimed to predict soil organic carbon, soil texture at several depths, as well as the thickness of the AB soil horizon and penetration resistance using a machine learning algorithm in combination with UAV images. We used an area in the Eurasian steppe zone (Republic of Bashkortostan, Russia) covered with the Stipa vegetation type as a test plot, and collected 192 soil samples from it. We estimated the models using a cross-validation approach and spatial prediction uncertainties. To improve the prediction performance, we also tested the inclusion of oblique geographic coordinates (OGCs) as covariates that reflect spatial position. The following results were achieved: (i) the predictive models demonstrated poor performance using only UAV images as predictors; (ii) the incorporation of OGCs slightly improved the predictions, whereas their uncertainties remained high. We conclude that the inability to accurately predict soil properties using these predictor variables (UAV and OGC) is likely due to the limited access to soil spectral signatures and the high variability of soil properties within what appears to be a homogeneous site, particularly in relation to soil-forming factors. Our results demonstrated the limitations of UAVs’ application for modeling soil properties on a site with homogeneous vegetation, whereas including spatial autocorrelation information can benefit and should be not ignored in further studies. Full article

Although ex situ conservation programs are increasingly valuable support tools for in situ conservation measures, success depends on these captive individuals to be genetically representative of the recipient population. The Eurasian stone-curlew (Burhinus oedicnemus) inhabit steppes that represent some of the most degraded and exploited habitats worldwide. A captive breeding program was implemented in Morocco as a pre-emptive effort for the conservation of the North African subspecies Burhinus oedicnemus saharae. However, the genetic origins of the founders of the captive flock were unknown. We applied a multi-locus approach to characterize the genetic ancestry of the current captive breeding flock by comparing it to wild populations from both Western and Eastern Morocco. Mitochondrial DNA and microsatellite markers were employed to assess levels of genetic diversity and relatedness within each sample, as well as potential genetic differentiation between wild and captive samples through PCA and admixture analyses. We recovered similar genetic diversity estimates, low levels of relatedness, and little differentiation between captive and wild samples. These results confirmed the Moroccan origin of the founders. We provide recommendations for the optimization of the Eurasian Stone Curlew conservation breeding program but also for future conservation breeding programs to ensure the effective conservation of genetic diversity and wild populations. Full article

Plant interference is a key factor influencing plant coexistence and species composition. The two primary forms of plant interference—allelopathy and resource competition—are often difficult to separate. This study conducted an outdoor pot experiment to quantify the distinct contributions of resource competition and allelopathy of Artemisia frigida on seedling growth of three species: Leymus chinensis, Cleistogenes squarrosa, and Potentilla acaulis. The index of relative neighbor effect (RNE) was used to quantify the overall effect of plant interference, while the inhibition rates (IRs) of resource competition and allelopathy were utilized to determine the specific contributions of allelopathy and resource competition from A. frigida on the growth of target plant species. The interference effect of A. frigida was found to be species-specific. The allelopathic effect of A. frigida played a major role in inhibiting the belowground biomass of L. chinensis (23.97%) and C. squarrosa (58.27%), while allelopathy and resource competition from A. frigida promoted the belowground biomass (45.12%) and aboveground biomass (46.63%) of P. acaulis, respectively. The combined effect of allelopathy and resource competition from A. frigida significantly affected the aboveground biomass of C. squarrosa and P. acaulis, as well as the belowground biomass of L. chinensis and C. squarrosa. These findings contribute to a better understanding of the patterns and mechanisms of plant species composition and its relationship with grazing intensity in this grassland ecosystem. Full article

As the most abundant group of mammals, rodents possess a very rich ecotype, which makes them ideal for studying the relationship between diet and host gut microecology. Zokors are specialized herbivorous rodents adapted to living underground. Unlike more generalized herbivorous rodents, they feed on the underground parts of grassland plants. There are two species of the genus Myospalax in the Eurasian steppes in China: one is Myospalax psilurus, which inhabits meadow grasslands and forest edge areas, and the other is M. aspalax, which inhabits typical grassland areas. How are the dietary choices of the two species adapted to long-term subterranean life, and what is the relationship of this diet with gut microbes? Are there unique indicator genera for their gut microbial communities? Relevant factors, such as the ability of both species to degrade cellulose, are not yet clear. In this study, we analyzed the gut bacterial communities and diet compositions of two species of zokors using 16S amplicon technology combined with macro-barcoding technology. We found that the diversity of gut microbial bacterial communities in M. psilurus was significantly higher than that in M. aspalax, and that the two species of zokors possessed different gut bacterial indicator genera. Differences in the feeding habits of the two species of zokors stem from food composition rather than diversity. Based on the results of Mantel analyses, the gut bacterial community of M. aspalax showed a significant positive correlation with the creeping-rooted type food, and there was a complementary relationship between the axis root-type-food- and the rhizome-type-food-dominated (containing bulb types and tuberous root types) food groups. Functional prediction based on KEGG found that M. psilurus possessed a stronger degradation ability in the same cellulose degradation pathway. Neutral modeling results show that the gut flora of the M. psilurus has a wider ecological niche compared to that of the M. aspalax. This provides a new perspective for understanding how rodents living underground in grassland areas respond to changes in food conditions. Full article

The grasslands in high-latitude areas are sensitive to climate warming and drought. However, the drought stress effect on the long-term variability of grassland productivity at the continental scale still hinders our understanding. Based on aboveground net primary production (ANPP) surveys, satellite remote sensing Normalized Difference Vegetation Index (NDVI), and meteorological data, we comprehensively analyzed three Aridity metrics and their effect on ANPP in Eurasian grassland from 1982 to 2020. Our results showed that the ANPP had an overall uptrend from 1982 to 2020, increasing most in the Tibetan Plateau alpine steppe subregion (TPSSR). Among three Aridity indicators, vapor pressure deficit (VPD) had an overall uptrend, while the trend of Aridity and soil moisture (SM) was insignificant from 1982 to 2020. Soil drought had negative effects on ANPP for all Eurasian grassland, while the atmospheric VPD had a positive effect on ANPP for TPSSR and the Mongolian Plateau steppe subregion (MPSSR), but a negative effect for the Black Sea–Kazakhstan steppe subregion (BKSSR) which was the driest subregion. SM had been the predominant driving factor for the interannual variability of ANPP in MPSSR since 1997. The increasing VPD had facilitated grassland productivity in alpine grasslands due to its cascading effect with an increasing temperature after 2000. The cascading effects networks of climate factors—drought factors (VPD, Aridity, and SM)—ANPP (CDA–CENet) indicated that SM was the predominant driving factor of the interannual variability of ANPP in MPSSR and BKSSR, and the dominance of SM had enhanced after the year 1997. The inhibitory effect of VPD on ANPP transformed into a facilitating effect after 1997, and the facilitating effect of SM is weakening in TPSSR. Full article

The Eurasian Iron Age Scythians, in all their regional iterations, are known for their lavish burials found in various kinds of tumuli. These tumuli, of varying sizes, are located throughout the Eurasian steppe. Based, at least partially, on the amounts and types of grave goods found within these mounds, the Scythians are usually modeled as militant, patriarchal mobile pastoralists, with rigid social structures. Yet, such interpretations are also due to accounts of Scythian lifeways provided by “classical” societies from the Greeks to the Persians, who saw the Scythians largely as barbarians, much like their neighbors to the north of the Greeks, the “Celts”. Despite recent interrogations of the barbarian trope, and the opportunity to dissect the classic formula of large mounds = elevated status, I contend that many studies on Scythian mortuary practices remain monolithic and under-theorized, especially by Western scholars. Drawing upon different conceptual and methodological frameworks, I present alternative, multi-scalar understandings of Scythian mortuary landscapes. Utilizing a spacetime-oriented, dialogical approach supplemented with geographic information systems, I interrogate how and why various meanings and experiences may have intersected in these protean Scythian landscapes of the dead, rather than reducing them to monolithic symbolic proxies of ideological status. Full article